I ran the first mission on the moon and only killed the hive with that weapon and as I was approaching to fill it up I slowed it down to kill them 1 by 1 till the upgrade notification popped up and as soon as it did I let them killed me and on the score screen it was exactly at 450 kills
Edit: if the actual kills needed it's 500 then I could guess that yellow bar enemies count for more since I killed a few of them
Looks like you'll get yours before I get mine. I would do the double bountie glitch to speed the process but I have so many other weapons to upgrade right now.
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