So during the GDC event they showed concept art of the original theming/design of the enemies. I noticed that there were 5 enemy races. Maybe I missed something, but I only know of the The VEX, The Fallen, The Cabal, and The Hive. Seems to be in that order left to right of the concept image. What's the fifth race? Are the triangles in the sky a reference to the pyrimidal spaceships they couldn't elaborate on during the panel?
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I'm going to go out on a limb here and say that fifth race is probably one of the nastiest. Assuming those giant onyx spaceships are theirs - which it appears to be judging by the triangle silhouettes in the OP art - in the concept art those ships look like they could straight up glass planets. Definitely a force to be reckoned with. Possibly even the race that originally decimated humanity to begin with.