Even though it isn't sourced on halopedian they do bring up how they are more like Rhinos. Bungie clarified that back in the day because people were calling them "space gorillas". I couldn't find anything in my brief look but i can dig something up if you really need to see.
i really dont give a damn what bungie says brutes look like or were modeled after. Bungie shouldve made them bald and given them ivory tusks if they wanted to remotely resemble a rhino [spoiler]i only work at a zoo. Not like i see these animals every day or anything[/spoiler]
It's their Hide. Just because they were modeled after something doesn't mean they have to look exactly like it. They were inspired by rhinos not brutes and that came from the devs themselves.