Guardians are the final shape, and humanity the victorious civilization. We are killing everything in site, and remaking the universe to our liking. Guardians can almost live forever. Remember, we were brought back to life by our ghosts, and are brought back every time we "die". Our golden age spread throughout the galaxy, and if real human history is any indication as to what we did when we were colonizing other worlds, maybe we are the darkness.
A galaxy has hundreds of billions of stars, we only spread through the Solar system in the Golden Age, which is essentially the worlds orbiting a single star, the sun. While we guardians are impressive, and we are great at killing, human civilization is not currently on par with civilizations that spread across countless star systems and timelines. Our Golden Age wasn't any more impressive than other other ages brought to other civilizations by the Traveler; the only difference was that the Traveler didn't get to flee like it always does because Rasputin crippled it, so it had to deploy the ghosts as an emergency measure.