We still don't know. If i were to guess i think PS4 and PS3 will be able to play and same for XB1 and XB360.
My logic: They would have to create servers for 4 separate worlds. Two of those worlds (PS3 and XB360) would be fading away fairly soon and the population of players would shrink. Since there is no battle between companies (Sony Vs MS)i think it would be a cost savings for both Bungie and Sony. Its just a guess. Maybe i'm just hopeful.
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Ya that makes sense, four servers would be challenging to manage, and the fact that Destiny is a minimum 10 year commitment for Bungie, having a couple servers that would just diminish over time seems pointless. I'm down with being hopeful, hopeful status just went up by 7%.