Find a slightly cheaper mobo and get a case that prioritizes cable management and you'll probably cut $125 right there
Try getting the OS for free... yeah good luck with that
You don't really need the optical drive since Steam's all downloads and even the newest Office Suite is all digital, but if you want one, get one that's cheaper than that, you don't need Blu-Ray
You'll only need the aftermarket cooler if you're overclocking, if not, scrap it
Edit: Doubt you'll need a gaming keyboard, too
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http://pcpartpicker.com/p/2d8KG Wargamer put this together for me... I'm new to the whole case and cable management thing and I've always preferred NVIDIA, but it seems quite solid to me.
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That's much better
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http://pcpartpicker.com/p/2d9Wk And here's mine (not really an AMD fan, but I should probably grab that 290)