Framerate, obviously. Despite the placebo effect, there is no discernible difference between 1080p and 4K resolution at the average seating distance for the average size tv. The difference that people see comes from the sharper picture UHD offers over HD. Resolution plays next to no part in it.
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What do you think makes the picture sharper, Einstein?
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Better tech that can display a sharper picture. If you look at the evolution of HD in gaming, it didn't even start out as sharp as it is now. The tech advanced, as well as knowledge of the platform, allowing for better graphical fidelity.
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So how does it display a sharper picture, unless there is, in fact, a sharper picture there for it to display? You're not thinking this through fully.
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Do I need to give you an example? Like, seriously? Even if you hate Destiny, play it on last gen then play the current gen version. Actually, you can do that with almost any game that released simultaneously across last and current gen. Current gen will have a sharper image and graphical fidelity even though they both exist on HD and not UHD. Why is that if they're both running on the same number of pixels? Because the better tech can display a better picture. Doesn't matter what the resolution is.
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Erm, the picture on the xbone is "sharper" than the one one the 360, [b]because it's a higher resolution[/b], dumbass. The 360 outputs through an upscaler, which works exactly the same way as the one in your TV does when you feed it SD content.