I bet Mara betrays us to the Witness.
We are relying heavily on her guidance and wisdom this season.
Who gave us the idea to make a stalemate with Xivu? Mara
Who has access to the vanaguard network where the Loki Crown Data is? Mara
Who has already been approached by the Witness? Mara
My prediction: Mara fires Rasputins arrays on the Traveler, wounding it, beginning the second Collapse.
Either it’s Mara or I’m just overlooking it could be Clovis 😂
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Like many people commented here already I don't believe Mara will be the one to betray us. Even in alternate timelines she has always been a steadfast ally (even if her methods are questionable at times) I think like you mentioned it might be more likely Clovis has some backup plan to betray us and become a disciple of the Witness in his grand scheme to become human's savior / emperor.