I recently read the Books of Sorrow and it's really got me thinking about stuff for Witch Queen and the upcoming seasons. I came up with this theory. It probably won't be true and I'm sure has holes aplenty but it's fun to think about.
So if you remember in Unraveling we learn about the whole thing with the Light and Darkness playing a game before reality existed, where flowers where put on an Infinite grid. The game always ended with one species of flower overtaking the others and preventing others from even existing. This went on until the Light for bored and put itself in the game as a factor leading to the Darkness following suit and creating reality in the process. Reality as a whole is essentially a continuation of this game but reading about Fundament makes me think that it is a more direct continuation of the game even if it's more on a metaphorical level. There's hundreds of sapient species living there and the hive eventually wipe all the others out.
Fundament seems to have been really important to the Light and Darkness because the Leviathan, a disciple of the Traveler, and the Worm God's, being that answer only to the Darkness itself, where both present on the planet. On top of that there's the Needle Ship. The description reminds me of a Pyramid Ship or a variant of it. We know the Traveler was also in the system helping out civilizations on Fundaments moons but that could just be normal Traveler stuff and not being there for anything specific.
But what really got me thinking was the part where the Hive God's encounter the Leviathan and it tries to convince them not to go find the Women Gods. It claims that the Krill/Proto Hive are it's proof against the Darkness. That they are it's hope that the Light's philosophy is true. I think that's why the Darkness was so intent on corrupting them. The Osmium King's familiar was a worm that guided the Hive God's to the worms and to me at least, it was implied that the Wom God's where the real reason for the Syzgy.
In Beyond Light the Darkness seemed to have a special interest in corrupting the Guardians to it's side for similar reasons. We where the champions of the Traveler and the Light so if it could get us to fight for it, then it would of struck a major blow against the Light. I think that the Traveler was likely going to give the Hive Ghosts once it finished up with the moons so that they could be champions of the Light like how the Guardians are now. We don't know if the Ghosts where actually created during the Collapse or if they existed before.
So when the Hive God's fell to darkness and took over Fundament, the Traveler no longer saw them as worthy of the Light. This could also explain why it never gave any other civilization access to the Ghosts. If the Hive, the very species that the Leviathan deemed as proof of the Lights philosophy could be corrupted, then it wouldn't want to take that risk again unless it could be completely sure. So when Savathûn inevitably exercises her worm and rejects the Darkness, the Traveler takes it as a sign of them being worthy again and gives Savathûn the light.
Like I said it's kind of a dumb theory but the idea of the Hive, one of the greatest threats in the Destiny universe, effectively being the original Guardians or at least coming close to it is a really interesting idea to me.
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It was never meant for the Krill/Proto Hive any more than it was for other creatures like the Eliksni and pre-Collapse Humanity. Ghosts were created as the Traveler's last ditch effort to prove a point to the Entity during the Collapse, and we happened to be the species it picked for this, so until we find a ghost or something similar that belongs to neither Humanity or Savathun's brood, then it's safe to say that we are the only race to legitimately "earn" the Traveler's favor. Savathun's ability to artificially create her own Hive Guardians, which only seem to work in her Throne World to our knowledge, is less of her being worthy of the Light and more of her ingenuity for managing to cheat both the Light and Dark at the same time.