I began having the Noah's Ark of connection errors a few nights ago, right after the update. Prior to that I had no issues. I had tried everything recommended, even though I did not believe it was related to my connection. In the end I changed everything back and had NAT open, standard DHCP connection with my Xbox 360.
Here's what fixed it (sorta).
At the time of the update, I had a level 7 titan. I had decided to try the hunter class, and created one in slot 2. I was able to play through the first tutorial level just fine. Once I went into the story mission 2 I could not stay connected for longer than a minute or two, most of the time disconnecting after launching to the surface, or immediately after it loaded and I was in the level. This also occurred if I tried to go to the tower.
Last night I decided to try playing with the titan after hours of trying to fix the connection issues and being dropped on the hunter. I was able to play without a single connection error for 2 hours, and never got an error. I played a story mission, a patrol for 45 min, visted the tower for quest stuff several times, and completed a strike in that time. Out of curiosity I tried to play the hunter at the end of my gaming session, and blam, connection error as soon as I landed on the surface.
So for now it appears I can play my titan character in slot one just fine, but if I try to play my hunter in slot 2 it's nothing but connection issues.
Hopefully this info helps you in fixing this issue.
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BUMP to see if anyone else finds it to be character related.